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If 'Add default .less theme' is checked in the New solution wizard, the properties.less and <solution_name>.less files are generated in the media node of the solution:

The properties.less file contains the theme properties, which are defined as less variables. The Servoy theme can be configured by changing the values of the properties defined in the properties.less file. NOTE CAREFULLY: you should change only the value of the defined variables. You should not remove any less variable or add any new less variable. 

The solution .less file may contain your own css/less styles and using variables from the properties.less file is allowed. Here you can also add your own .less variables which can be used in your own less style.